Photo of the day (126): Epiphragmophora

This picture was made by Lucio Spiderman in Argentina, Jujuy, Yungas district, Peña Alta. The snail is probably Epiphragmophora cryptomphala Ancey, 1897. 

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Wood & Gallichan (2008: 39) wrote that Ancey spelled the name as eryptomphala in the description and cryptomphala in the plate legend of the same paper. “We are unaware of any published corrections of eryptomphala, but it almost certainly an original incorrect spelling. The correct original spelling should be fixed by the First Reviser if this has not already been achieved”.
The revision by Cuezzo (2006) did not mention the spelling difference in the original publication, but she uses the name E. cryptomphala.

There are two other species which may be confused with E. cryptomphala: E. trigrammephora d’Orbigny, 1835 and E. walshi Cuezzo, 2006. Cuezzo (2006: 132) writes “Parodiz (1957) and Fernández (1973) considered E. cryptomphala to be a subspecies of E. trigrammephora. However, the main differences between E. cryptomphala and E. trigrammephora are found in both the shell and genitalia. In E. cryptomphalathe shell umbilicus is completely overlapped or fused with the columelar side ofthe peristome. The shape of the aperture is not as constant as in E. trigrammephorabeing subquadrangular to subovoidal, while in E. trigrammephora, all shells examined have a subquadrangular aperture.”. And “Epiphragmophora cryptomphala is similar to E. walshi n. sp. in that both have three peripheral bands in the shell. Also, both species live in Salta Province and are typical from xerophilic environments. Epiphragmophora cryptomphala differs from E. walshi n. sp. in the umbilicus that is narrower in E. walshi n. sp. and not overlapped by peristomal basal fold. Epiphragmophora walshi n. sp. is smaller in shell diameter than E. cryptomphala. Differences in the genitalia are mainly in the shape of mucous glands, insertion of their efferent ducts and shape and length of dart sac”.
The differences are thus subtle and as usually an expert eye may give a different opinion. The following picture are copied from Cuezzo (2006: Fig. 5D E. trigrammephora, Fig. 2A E. cryptomphala, Fig. 7C E. walshi; note that the scale bar does not apply to all three figures).

Thanks Lucio for sharing this photograph.

References:
Cuezzo, M.G., 2006. Systematic revision and cladistic analysis of Epiphragmophora Doering from Argentina and southern Bolivia (Gastropoda, Stylommatophora: Xanthonychidae). – Malacologia 49: 121-188.
Wood, H. & Gallichan, J., 2008. The new molluscan names of César-Marie-Felix Ancey including illustrated type material from the National Museum of Wales. Biotir Reports (Cardiff) 3: i-vi, 1-162.

 

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