Already some months ago, Thomas Watters published a new revision dealing with Hispaniolan annulariids (Watters & Larson, 2017).
“The annulariid genera Chondropomella, Chondropomium, and Clydonopoma are believed to have originated from the Tiburon/Barahona Peninsula in isolation from the rest of Hispaniola. Chondropomium has colonized the rift valley and adjacent river valleys between the Tiburon Peninsula and the remainder of Hispaniola. It is primarily limited to xeric lowlands, rarely found above 200 m elevation. Little is known about the rare Chondropomella but they seem to occur in the rift valley in xeric areas as well. In contrast, Clydonopoma is endemic to the Sierra Baoruco with a single species in the adjacent eastern Massif de la Selle and occupies the upland mesic forests and pine savannahs between 200–4000 m. The most widely distributed species, Chondropomium weinlandi, has been the subject of considerable confusion concerning the nature of its many color forms or subspecies, as well as its valid name. This species was investigated using phylogenetic methods and compared to congeners and related genera. A phylogenetic study aimed at elucidating relationships among these taxa analyzed a partitioned matrix of nuclear (ITS 1) and mitochondrial (CO1, 12S, 16S) DNA sequences in a Maximum-likelihood framework under the GTR+G substitution model. Contrary to Bartsch’s 1946 assessment that C. weinlandi is a complex of subspecies, it is here shown to be a single, highly polymorphic species for color. The new genus Superbipoma is recognized based on phylogenetic, radular, and conchological evidence. It contains two species: S. asymmetricum (Henderson and Simpson, 1902) and S. superbum (Pilsbry, 1933). Eleven species of Chondropomium are recognized including two new species: C. caelicum and C. sardonyx; three species of Chondropomella and nine species of Clydonopoma, including one new species, Clydonopoma titanum, are recognized. A calcified operculum is the ancestral condition for the Annulariidae. In Clydonopoma and Chondropomella the operculum is a particularly complex calcified structure termed the pseudolamella. This structure has been lost in Chondropomium and Superbipoma”.
As usual, this paper is very thorough, well organised and includes a phylogenetic study based on a multi-locus analysis.
Watters, G.T. & Larson, P., 2017. A revision of the Hispaniolan genera Chondropomella, Chondropomium, and Clydonopoma (Gastropoda: Annulariidae), with the recognition of a new genus, Superbipoma: phylogenetic, radular, and conchological evidence. – Nautilus, 131 (3):163-201.